| 
    
        
            | PRACTICAL ARABLE WILD FLOWER
                CONSERVATION |  ...Practical Arable Wild Flower Conservation on the
      Kemerton Estate - By J Clarke & C C Morgan July 2000 SummaryKemerton Estate is in south Worcestershire – on
the flanks of Bredon Hill – on soils varying from Limestone Brash, through heavy clays to sand and gravel soils and
valley silt. Kemerton Conservation Trust (KCT) is a charity set up to promote nature conservation in the Bredon
Hill area. It advises the estate on conservation matters and manages specific conservation
sites.
 Other research at Kemerton and elsewhere has focussed on benefits for wildlife and costs of
      arable margins. This paper concentrates on the practical implications of establishing new sites and enhancing
      the botanical richness and diversity of existing sites. The following points were
      considered: 
 
    
        
            
                | We found that arable wild flower
                      conservation margins were more easily established on sites with a basic, existing arable
                      flora and without pernicious grass weeds. Kemerton follows a policy of using, whenever
                      possible, seed sourced from within the estate boundaries. To achieve this and to obtain
                      sufficient seed, KCT operates a wild flower nursery and has found that the period between
                      initial seed collection and final sowing/planting covers from three to four years. It is
                      important that seed collection, seed batches and later destinations are accurately recorded.
                      It is also important to obtain accurate baseline data as well as ongoing monitoring of the
                      project. Arable wild flower conservation margins may require additional management to control
                      pernicious weeds or to clean the crop. To date it appears that soil types (i.e. fertility)
                      may influence the amount of management required to control pernicious weeds. Success rates
                      for sowing direct and planting plugs were similar. The use of ‘cages’ to protect
                      ‘introductions’ is important. Costs of growing or purchasing seed will govern sowing rates
                      but we aim to sow as much as possible. |  Oathill Arable Margin
 |  IntroductionThe Kemerton Estate is owned by the Darby Family and is managed by Adrian
      Darby, formerly chairman of the Royal Society for the Protection of Birds and currently chairman of
      Plantlife. Since the early eighties it has tried to integrate farming and nature conservation and to set an
      example to other landowners and farmers. The estate comprises some 950 acres and is situated on the southwest
      slopes of Bredon Hill in Worcestershire, overlooking the Avon and Severn Valleys. It contains a diverse range
      of habitats and species.  These have been maintained and improved through sympathetic management of the
      farmed areas and by the establishment of specific estate ‘nature reserves’.
 Since the early
eighties, a full time conservation officer has been employed to manage conservation sites, monitor the wildlife of
the estate and to work closely with the farm manager, P. Doble. This post is funded by the Kemerton Conservation
Trust, a charity established by Mr Darby in  1989 to promote nature conservation on the estate and its
surroundings. AimsTo investigate practical ways of enhancing existing ‘rich’ arable wild flower conservation margins and of
establishing new sites.
 BackgroundMuch research has already carried out by the Game Conservancy Trust and others into the effects on wildlife of
arable conservation margins.
 An earlier Conservation Officer at Kemerton, (Deane 1989), combined a great deal of earlier
      research to produce a report which described ‘expanded field margins’, including arable wild flower margins –
      comparing costs involved and benefits to wildlife. Having accepted that much is to be gained by managing these margins, when establishing new sites
      and enhancing the botanical richness and diversity of existing sites the following questions were
      considered: 1 Selection of
MarginsRobert Deane surveyed most arable margins on the estate and identified two on Limestone Brash that were
particularly rich. Over the next few years, work continued on managing (farming) and monitoring the flora of these
sites.
 In 1997, as part of the restoration of a gravel pit on the estate into a nature reserve, a one
      hectare area that was unable to be quarried, due to the archaeological interest underneath was restored as a
      spring sown arable area, which is not harvested but left as a food source for wintering
      birds. In 1998 two six-metre margins on wet, clay ground (Runway and Lakeside) were selected as
      possible arable conservation wild flower margins. They were chosen as they were on a different soil type and
      because there had been historical records for Corn Buttercup Ranunculus arvensis and both Sharp–leaved Fluellen Kickxia elatine and Round-leaved Fluellen Kickxia spuria in Lakeside. In 1999 Lakeside continued but as a 1/3 hectare
      sown wild bird cover under Set-aside. In 1999 a 1/3 hectare headland on lighter soil – in Chapel field – was also selected under
      Set-aside. A range of arable wild flowers had been identified on the overlapping ground between a
      wet-grassland conservation area and the arable crop. The records included Petty Spurge Euphorbia peplus, Corn Mint Mentha
      arvensis and K. spuria. In 1999 a six-metre margin on sandy soils in Welsh Furlong was selected to be investigated and
      was left unsprayed. The margin is adjacent to a busy public footpath which is annually topped so does not
      have a Black-grass Alopecurus myosuroides
      problem. This margin is currently under a two-year investigation. To date although no rarities have
      occurred, it has a good, basic flora with potential for enhancement. As with the limestone headlands,
      numerous favourable comments have been received from people using the footpath. In 2000 three further six-metre margins on the Limestone Brash and adjacent to the existing
      arable wild flower conservation margins, were selected under a Whole Estate Countryside Stewardship
      agreement.   2 ProvenanceThe Trust believes that seed should be sourced as close as
      possible to its intended destination. This retains the genetic integrity. At Kemerton, wherever possible seed
      is taken from the estate. However, for Shepherd’s Needle Scandix
      pecten – veneris, Corn buttercup Ranunculus arvensis and Field
      Gromwell Lithospermum arvense our nearest source is within ten miles - near Pershore. Corn Salad
      Valerianella locusta and V.
      dentata have been collected elsewhere on Bredon Hill.
 3 Obtaining
Sufficient Seed At Kemerton we have found that the viability and
germination rate of arable wild flowers is unpredictable and therefore it is better to have as much seed as is
realistically possible. To achieve this, the Trust takes seed and grows the plants in ‘controlled conditions’. For
many plants a small patch of ground is used as an arable wild flower nursery. The seeds of these species are easier
to harvest. It must also be considered that most arable plants flower and seed over a long period and harvesting
will require numerous, time-consuming visits. Poppy Papaver species shed
seed when the seed pod is shaken and therefore must be harvested more frequently in windy conditions. When
harvesting seedpods of Night-flowering Catchfly Silene noctiflora it is
important to check for the presence of larvae of moths such as Hadena
confusa and Hadena bicruri which feed on the seeds of Silene species! Some plant
species, such as V. locusta, Venus’ Looking Glass Legousia hybrida and Scarlet Pimpernel Anagallis arvensis readily drop their small seeds. These are grown in pots or
trays in a cold greenhouse. When flowering begins the pots are stood on sheets of paper that catch the seeds as
they fall. Every few days, the seeds can then be easily collected The collected seed is stored carefully in
individually labelled bags, within metal containers in cool conditions.
 Some arable wild flowers germinate in autumn, some winter/spring and others in spring (Fitter
      & Attenborough, 1987) – and this must be born in mind when planning planting. When planning a scheme of this type it must be noted that it may take three to four years to
      before sufficient seed has been produced. At Kemerton, the plants have been introduced into sites both as
      seed and as ‘plugs’. 4
RecordingEach batch of seed collected
from the wild is allocated a code number. This comprises a number for the year, the Latin initials of the species
and collection number. For example 98/PA/1 is the code number for seed from Papaver argemone, collected in 1998 – from a particular site. Further details are
recorded on computer spreadsheets. Subsequent seed collections from the original 98/PA/1 will be allocated a
related code (e.g. 99/PA/1) and entered on computer. Example:
 
    
        
            
                
                    | Scientific Name | Common Name | Provenance | Date | Collector | Germination | Sown Where | Code No. | Planted Out |  
                    | Papaver argemone | Prickly Poppy | Oathill | 5/7/98 | JC | Spring | pot1/2/99 | 98/PA/1 | Chapel Field |  The spreadsheet also contains a
      space for notes. If Papaver argemone was also collected from a second
      site in 1998, then the code number allocated would be 98/PA/2. 5 Monitoring a.  Existing SitesAs part of a ten-year monitoring strategy for the estate, these non-Stewardship margins are intensively surveyed in
years 1,6 & 10. These intensive surveys are supported by annual casual observations.
 The method used for intensive survey will be
      a fixed transect along the complete length of the margin, with a 1 metre quadrat 1 metre in from the edge,
      every 50 metres along the transect line. Presence and  % cover will be recorded for each species within
      the quadrat. b. New/Stewardship
      Sites These will be monitored as above, but more frequently.
 This work is part of an estate-wide
      monitoring strategy supported under English Nature’s ‘Biodiversity Grant Scheme’. 6 Management a. Kemerton
      Lake  – Nature Reserve – sandy soilsThe site is spring-sown with a mixture of Linseed, cereal and Quinoa as wild bird winter feed. In addition, for two
years attempts have been made to ‘bulk up’ the existing diverse arable wild flower population and to introduce
other desirable species. Aside from any direct wildlife benefits, this ‘reservoir’ of Kemerton plants has two other
functions – to act as a ‘reserve’ population should disaster occur elsewhere, and to provide a demonstration area
for visitors.
 As the site is on a nature reserve, several problems have been identified.   Sowing dates usually follow on from the main farm spring sowings,
      thus providing later grazing for pigeons and rabbits. The site supports a large rabbit population. Efforts
      are being made to fence out and cull the rabbits. It is interesting to note that rabbits appear to locate and
      selectively feed on the native, introduced plants. In addition, up to 50 Canada Geese attempt to breed in or
      near the reserve and heavily graze the arable area. In future culling will take place under
      licence. Attempts to protect seed and plants of introduced arable wild flowers by use of individual wire
      cages have been partially successful – geese can reach inside and rabbits and hares may pull them over. An
      improved design trialled in Chapel field (see below) will be used in the future. Common Couch Elymus repens is a problem on the site. The method that will be trialled
      later this year is spraying of ‘ Roundup’ on the affected area once all the cover crops and arable
      wildflowers have set seed. This will kill all the plants, including the E. repens, but leave unaffected the shed arable wildflower seed and the
      seed on the crop which will be eaten by the wintering birds. b.  
      Chapel Field – heavy soilsIn autumn 1999 this headland was sown with a ‘Wild Bird Cover Crop’ under set-aside and will therefore not be
harvested.  A light dressing of fertiliser is permitted. A.
myosuroides was not sprayed in 2000 but will need to be controlled in the future.  As mentioned earlier,
this site had already been identified as a ‘good’ arable wild flower site. Trials are taking place to introduce
further suitable species.  These have been planted as seed and/or plugs in 2m square wire exclosures, ½ metre
high - low enough to avoid the spray boom.  This method has proved hugely successful by eliminating grazing
problems and in providing a favourable microclimate for germination and growth.  To protect any genetic
integrity, it was decided that should any introduced species be found outside the cages during the first year, then
the introduced plants would be destroyed.  This cage design will be used on other sites.  Chapel Field is
proving to be an important arable wild flower area on the heavier soils.
 c.   
      Runway Field and Lakeside – Heavy claysIn autumn 1998, these headlands were planted with wild flower seed.  In spring 1999 wild flower plugs were
planted.  The extreme soil conditions in this wet clay and rabbit grazing resulted in apparent complete
failure.  In autumn 1999 a further attempt was made in Lakeside, using the larger cages (as described for
Chapel Field).   This has been a partial success – Scandix pecten
- veneris and R. arvensis  thrived best.  In addition,
several plants of the former were found outside the exclosures – probably seed from 1998. A. myosuroides was not sprayed in 2000 but will need to be controlled in the
future.  Lakeside would appear to have limited potential but trials should continue.
 d.  
      Welsh Furlong – Sands and siltsThis trial headland is not included in any
      current conservation or agri-scheme.  Because of its possible floristic interest no species
      introductions will be tried until the end of a two-year monitoring period.
 e.  
      Stewardship Sites – Limestone brashExisting (pre-2000) sites – Oathill North and Westmancote Piece East.
 These headlands had been identified by Deane
      (1989) and have been managed for their arable wild flower interest since then.  Indeed, they have been
      the main seed source for the arable wild flower project at Kemerton.  They are sited next to grass
      margins/footpaths, and invading grass has been a localised problem but can be controlled with sprays. 
      Some of the seed and plants produced in the nursery have been planted back into the headland – to ‘bulk up’
      the existing seed bank.  Apart from regular monitoring it is unlikely that further management will be
      required.  These headlands are used as demonstration sites for visitors to Kemerton
      Estate. New (post-2000) sites – Westmancote Piece South, Westmancote Piece North, Oathill East and Pad
      Piece North – Limestone brash. Because of the botanical importance of the neighbouring arable headlands, the opportunity to
      increase the number of conservation headlands was taken under the Countryside Stewardship Scheme. 
      Kemerton Countryside Stewardship margins are different from ‘normal’ Countryside Stewardship margins as the
      headland can be cropped. Once the monitoring programme has assessed their floristic value, a decision on
      whether to carry out ‘bulking up’ or an introduction programme will be made.  From experience gained on
      the existing sites, it is likely that some control of invasive grasses will be necessary.
        Conclusions/RecommendationsIt is not always easy to produce a good arable conservation margin – particularly on heavy/wet ground or where few
arable wild flower species occur naturally.  Generally, a soil of low fertility will ensure that the site is
not dominated by pernicious weeds and that the arable wildflowers do not compete too much with the crop. (P. Doble
pers. com.)
 Broad-leaved crops such as brassicas and beans in a crop rotation provide a good opportunity to
      control A. myosuroides and Wild Oat Avena fatua using graminicides whilst leaving arable wildflowers relatively
      unaffected.  However, graminicides sometimes seem to affect wildflower seedlings (P.Doble. pers.
      com.) Grass adjacent to arable wildflower conservation headlands should be regularly cut to prevent
      the grass seeding into the headland. Spring-sown cereals tend to have a less dense canopy than autumn sown crops.  However, some
      arable wildflower seeds germinate in the autumn. Rabbits appear to selectively graze ‘native’ plants in preference to planted
      crops. Where rabbit populations are high, establishing an arable wildflower conservation margin may be
      very difficult. Growing arable wildflowers in nursery or greenhouse conditions is labour-intensive. 
      However, harvesting is easy and the maximum quantity of seed can be collected. Our results show that success rates for planting seeds directly into newly sown headlands and
      planting out pot grown plugs are similar.  Therefore in most circumstances it is preferable to sow
      direct. The use of cages to protect seedlings during the first year or two of introductions is
      preferred, however only small sections need protection. It is not possible to recommend sowing rates.  However, to avoid repeating the process it
      is better to sow as much seed as is practical and affordable. At Kemerton it is recommended that suitable sites for arable wild flower conservation margins be
      eventually established on examples of all soil types.  The area available for conservation will always
      depend on current farm practice and policy.  Meanwhile, in the next few years trials should take place
      on sites that still retain some interesting plant species, or from where past records indicate potential.
      Sites already identified include a section of Runway Field (old record of Cornflower Centaurea cyanus), other sections of Lakeside, Deer Field (80’s records of
      R. arvensis and both Kickxia spp.), Butchers, Long Ground, and Welsh Furlong (already being
      investigated). Ideally, other farmers should be encouraged to follow Kemerton’s example regarding the sourcing
      and growing of wild flower seeds.  As arable wildflower conservation margins are now part of Countryside
      Stewardship and (indirectly) affected by Set-aside rules, additional financial support should be made
      available for farmers who retain the local genetic integrity of arable wild flowers. Some farmers may not have the resources to
      collect and grow seed in the way that Kemerton does - nor may they be aware of local seed sources. Therefore
      both local and centralised information is required.  This could be compiled and held at national level
      by Plantlife, Flora Locale or at local level, by FWAG.  Additionally, a network of regionally appointed
      nurseries could supply seed of local provenance.   
 AcknowledgementsThanks to A.M.G.Darby and P.Doble for comments on the early draft and M.J.Morgan for reading final
draft.
 
 Bibliography 
    
        Andrews J. Rebane M. (1994) Farming and Wildlife. A Practical Management Handbook. RSPB,
              Bedfordshire
        Bruce B. (1999) Biodiversity Action Plan for Worcestershire. Worcestershire Wildlife
              Trust, Worcestershire
        Clarke J. Clarke P.(1987) reconciling farming with wildlife. A case study of
              conservation management techniques on Kemerton Farm 1983-1987.Kemerton Trustees Ltd,
              Gloucestershire
        Clarke J. Clarke P. Offer D. (1995) Nature Conservation Plan. Kemerton Estate. Kemerton
              Trustees Ltd, Gloucestershire
        Clarke J. Morgan C. (2000) Kemerton Estate Monitoring Strategy. Kemerton Conservation
              Trust, Gloucestershire
        Deane R. J. L. (1989) Expanded Field Margins. Their costs to the farmer and benefits to
              wildlife. A report to the Nature Conservancy Council, Kemerton Court,
              Gloucestershire
        Fitter A. Attenborough D (ed). (1987) Collins New Generation Guide to the Wild Flowers
              of Britain and Northern Europe. Collins, London
        Rose F. (1981) The Wildflower Key. Warne, London
        Close to the Edge. (March 2000) English Nature Magazine
        No 48.English Nature. Peterborough 
 Appendix 1
                            
Kemerton Arable Wildflower Trials
 a.  Aims - To ‘Bulk Up’
Existing Conservation Sites
 Year One Collect Seed From
   Existing SiteLabel and Record
 Year Two Plant Seeds in
   ‘Controlled Environment’Sow ‘Fine’ Seeds in Greenhouse in trays and in plugs
 Plant out Plugs in Headlands - in Spring - Record
 Harvest Seed from Greenhouse and Nursery in Summer – Label and Record
 Year ThreeSow some (or all) Seed Direct into
   Headlands
 Year FourSow Direct Into Headlands
 
 b.   Aims - To Introduce/Re-introduce Arable Plant Species into
      New/PreviousSites Year OneCollect Seed From Nearest Local Source - Label and Record
 Year TwoPlant Seeds in ‘Controlled Environment’
 Sow ‘Coarse’ Seeds in Nursery and some in Greenhouse as Plugs
 Plant out Plugs in Headlands - in Spring - Record
 Harvest Seed from Greenhouse and Nursery in Summer – Label and Record
 Year ThreeIf more Work is Required  - Repeat Year Two
 
 Year Four
 Sow Direct Into Headlands
 
 Appendix 2      Arable Wildflower Species List For Kemerton
      Estate 
    
        
            | Scientific Name | Common Name |  
            | Aethusa
                cynapium    | Fool’s Parsley |  
            | Agrostemme
                githago    | Corn Cockle (introduced)   |  
            | Alliaria
                petiolata    | Garlic Mustard |  
            | Alopecurus
                myosuroides    | Black Grass |  
            | Anagallia
                arvensis    | Scarlet Pimpernel |  
            | Aphanes arvensis    | Parsley Piert |  
            | Atriplex patula    | Common Orache |  
            | Barbarea
                vulgaris | Wintercress |  
            | Brassica
                nigra | Black Mustard
                  |  
            | Bromus
                sterilis | Sterile Brome |  
            | Capsella
                bursa-pastoris | Shepherd’s Purse |  
            | Centaurea
                cyanus | Cornflower |  
            | Cerastium
                fontanum | Common Mouse-ear |  
            | Chaenorhinum
                minus | Small Toadflax |  
            | Chenopodium
                album | Fat Hen |  
            | Cirsium
                arvense | Creeping Thistle |  
            | Conium
                maculatum | Hemlock |  
            | Coronopus
                didymus | Lesser Swinecress |  
            | Coronopus
                squamatus | Swinecress |  
            | Dactylis
                glomerata | Cocksfoot Grass |  
            | Euphorbia
                exigua | Small Spurge |  
            | Euphorbia
                helioscopia | Sun Spurge |  
            | Euphorbia
                peplus | Petty Spurge |  
            | Fallopia
                convolvulus | Black Bindweed |  
            | Fumaria
                officianalis | Common Fumitory |  
            | Galium
                aparine | Cleavers |  
            | Geranium
                dissectum | Cut-leaved Cranesbill |  
            | Geranium
                molle | Dove’s Foot Cranesbill |  
            | Hypericum
                perforatum | Perforate St.John’s Wort |  
            | Kickxia
                elatine | Sharp-leaved Fluellen |  
            | Kickxia
                spuria | Round-leaved Fluellen |  
            | Lamium
                amplexicaule | Hen-bit Deadnettle |  
            | Lamium
                purpureum | Red Dead-nettle |  
            | Legousia
                hybrida | Venus’ Looking Glass |  
            | Lepidum
                campestre | Field Pepperwort |  
            | Lithospermum
                arvense | Field Gromwell (introduced) |  
            | Matricaria
                matricarioides | Pineappleweed |  
            | Matricaria
                recutita | Scented Mayweed |  
            | Medicago
                lupulina | Black Medick |  
            | Mentha
                arvensis | Corn Mint |  
            | Papaver
                argemone | Prickly Poppy |  
            | Papaver
                dubium | Long-headed Poppy |  
            | Papaver
                hybridum | Rough Poppy |  
            | Papaver
                lecoqii | Babbington’s Poppy |  
            | Papaver
                rhoeas | Common Poppy |  
            | Persicaria
                maculosa | Redshank |  
            | Ranaunculus
                arvensis | Corn Buttercup(re-introduced) |  
            | Raphanus
                raphanistrum | Wild Radish |  
            | Reseda
                lutea | Wild Mignonette |  
            | Reseda
                luteola | Weld |  
            | Scandix
                pecten-veneris | Shepherd’s Needle(introduced) |  
            | Sheradia
                arvensis | Field Madder |  
            | Silene
                alba | White Campion |  
            | Silene
                noctiflora | Night-flowering Catchfly |  
            | Sinapsis
                arvensis | Charlock |  
            | Sison
                amomum | Stone Parsley |  
            | Sonchus
                asper | Prickly Sow Thistle |  
            | Stellaria
                media | Common Chickweed |  
            | Thlapsi
                arvense | Field Pennycress |  
            | Triplospermum
                inodorum | Scentless Mayweed |  
            | Urtica
                dioica | Common Nettle |  
            | Urtica
                urens | Small Nettle |  
            | Valerianella
                dentata | Narrow-fruited Corn Salad(introduced) |  
            | Valerianella
                locusta | Corn Salad (introduced) |  
            | Veronica
                agrestis | Green Field Speedwell |  
            | Veronica
                arvensis | Wall Speedwell |  
            | Veronica
                persica | Field Speedwell |  
            | Veronica
                polita | Grey Field Speedwell |  
            | Viola
                arvensis | Field Pansy |  
            | Viola
                tricolor | Wild Pansy |    |